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Extensive scientific research documents that vegetated strips of land along waterways provide extensive water quality and other environmental and economic benfits.
Scientific research clearly documents that riparian buffers, particularly forested buffers and those along headwater streams, deliver tremendous benefits. Through the interaction of their soils, hydrology, and biotic communities, riparian buffers serve many important physical, biological, and ecological functions (Klapproth, 2009).
Riparian buffers are the lands and assemblages of plants bordering rivers, streams, bays and other waterways. They directly affect and are directly impacted by the aquatic environment. Buffers have high levels of soil moisture, experience frequent flooding, and are populated by plant and animal communities that are adapted to life along the water. The boundary between the buffer and adjoining uplands is gradual and may not be well defined (Klapproth, 2009).
The USDA Forest Service estimates that over one-third of the rivers and streams in Pennsylvania have had their riparian buffers degraded or altered, a sobering statistic when the value of their functions is considered (DEP, 2006).
Scientific research clearly documents that riparian buffers, particularly forested buffers and those along headwater streams, deliver tremendous economic, ecological and other benefits. Among these benefits, riparian buffers:
The width needed for a riparian buffer to be effective depends on a number of factors, but, in general, the wider the buffer, the greater the benefits delivered.
Forested riparian buffers provide substantially more and better ecosystem services than grass buffers (Burgess, 2004). The roots of herbaceous and woody plants strengthen the stream bank and prevent stream bank erosion. Roots and downed trees slow the flow of stormwater and form a physical barrier to the stream or river, which allows sediment to settle out and be trapped. The forest canopy shades water, moderating water temperature. The plants are an important source of woody material in streams, which provides habitat and food for aquatic wildlife. They also provide quality habitat and food for terrestrial wildlife. These services are discussed in detail below.
As described in the Headwater Streams section below, research demonstrates that healthy riparian buffers along headwaters streams, both perennial and intermittent, deliver exceptionally high ecological value.
The following sections highlight key ecosystem services delivered by riparian buffers:
Riparian buffers reduce erosion, which both conserves topsoil and lessens the amount of sediment in streams and rivers. A buffer’s roots of herbaceous and woody plants strengthen the stream bank by going through the topsoil and into a stream bank’s weathered or fractured bedrock and other more stable strata. This increases the stream bank cohesiveness and adds a tensile strength that can resist shear stresses on stream bank soil (Castelle, 2000).
Riparian buffers filter sediment from stormwater runoff, reducing the amount of sediment in streams and rivers. Tree roots and downed trees slow the flow of surface water and form a physical barrier, which allows sediment to settle out and be trapped. Several studies have shown the effectiveness of riparian buffers in filtering sediment, including:
Filtration of sediment is also important for removing chemical pollutants that bind to sediment. For example, excess phosphorus binds to soil and is found primarily in the top few inches of the soil, which are very susceptible to erosion. Trapping sediments is the most effective way to reduce non-point source pollution (Bongard, 2009).
Riparian vegetation removes metals, nutrients, and other chemicals from runoff via plant uptake and by facilitating bacterial degradation of the pollutants (Castelle & Johnson, 2000 Although narrow buffers can generally remove sediment in runoff, wide buffers are needed for effective nutrient removal (Dabney, Moore, & Locke, 2006).
The removal of nitrogen, a major pollutant of many watersheds, from runoff occurs almost exclusively in water-saturated zones where abundant organic matter is present. Bacteria in the buffer use nitrogen as an energy source, converting it to gas. Plant roots also absorb nitrogen in groundwater and use it for plant growth. Buffers act as a nitrogen sink when it is taken up by trees and stored in their biomass.
Multiple studies have shown that buffers are effective in removing pollutants from water:
The trees of riparian buffers shade the water, moderating water temperature. Temperature is a critical influence in aquatic ecosystems, affecting both the physical and biological characteristics of the stream. Changes in temperature can decrease stream biodiversity and impede animal growth. Increases in summer temperatures can increase the susceptibility of fish to pathogens; decrease food availability; alter the feeding activity and body metabolism of fish; inhibit spawning, and block spawning runs into streams (Castelle and Johnson, 2000). At the same time higher stream temperatures reduce the amount of dissolved oxygen in water; they also increase the metabolic rate of aquatic animals, increasing their oxygen needs.
In small streams, the presence of a forest canopy greatly affects the intensity of light reaching the surface of the stream. Depending on the season, light intensity in a shaded area of a stream can be 30 to 60% less than that of an exposed area (Sweeney, 1992). By limiting the amount of solar radiation that can reach a stream, trees limit both the daily fluctuations in stream temperature and the maximum stream temperatures reached (Bongard, 2009). A British Columbia study found that streams without buffers have temperatures up to 1-2 oC higher than those with buffers (Rayne, Henderson, Gill, & Forest, 2008). A study from Washington State found that non-buffered streams have maximum temperatures 2.4 oC higher than those with buffers (Pollock, Beechie, Liermann, & Bigley, 2009). In Oregon, studies of stream temperatures following the removal of riparian vegetation found that maximum stream temperatures both increased by 7 °C and occurred earlier in the summer. (Shifts in the timing of maximum temperatures, with greater increases in early summer stream temperatures, can impact sensitive stages of aquatic animals.)
Increased water temperature increases the toxicity of many chemicals, such as ammonia. Ammonia is an inorganic form of nitrogen. It is present in water in two forms, un-ionized (NH3), which has a relatively high toxicity, and ionized (NH4+), which has a relatively negligible toxicity. As water temperatures increase, more of the ammonia is converted to the toxic un-ionized ammonia form (EPA, 1995). Polluted runoff is a large source of ammonia and nitrogen to streams (EPA, 1995). When riparian buffers are not preserved, both their ability to remove nitrogen from runoff and their ability to maintain lower water temperatures and prevent it from converting to its un-ionized ammonia form are lost.
Large woody debris is an essential part of stream life. It provides fish habitat and changes the stream’s physical condition. Organic matter from riparian buffers, such as leaves, twigs, logs and stems that fall from the buffer into the water are a main source of food for aquatic macroinvertebrates. Aquatic macroinvertebrates are animals without a backbone, are visible with the naked eye and spend all or part of their life in the water. These animals, which include worms, mollusks, insects and crustaceans, consume the wood and the biofilms (bacteria, fungi, and algae) that form on it (Pitt & Batzer, 2011), serving as a vital link in the food web between the producers (e.g. leaves, algae) and higher consumers, such as fish.
The wood from buffers also traps additional leaf litter and wood. Macroinvertebrates use the wood as habitat, living inside the wood, under residual bark, and on surfaces that protrude out of the water. Some insects use the protruding surfaces as sites to emerge into adults or to lay eggs (Pitt & Batzer, 2011). A study of 16 streams in eastern Pennsylvania found that forested stream segments have over six times the amount of large woody debris than do grass buffered streams, even though two-thirds of the grass buffered streams were immediately downstream of forested areas (Sweeney, 1992)
Forested riparian buffers are also essential for maintaining stream and river bottom habitat. Most of the biological activity in stream ecosystems takes place on inorganic (sand, gravel, cobble, etc.) and organic (leaves, woody debris, etc.) materials on stream bottoms. Networks of tree roots, the organic debris from buffers and the variety of sizes of cobble and gravel these trap can increase the overall size of bottom habitat more than a thousand times when compared to a bare mineral soil bottom in a grass-buffered stream (Sweeney, 1992). In addition, where riparian buffers have been deforested, streams are narrower because of encroachment by herbaceous plants, mostly grasses, that would have been shaded out under forest cover, causing an additional loss of river bottom habitat (Sweeney, 1992).
Deforestation of a section of a riparian buffer can change stream bottom habitat and influence biodiversity, even if the deforested section is still vegetated. In southern Appalachia, 12 streams with deforested, but vegetated, buffers were studied. The deforested sections were up to 5.3 km long. The stream segments studied were all downslope of watersheds with at least 95% forest cover. As the length of deforested sections increased, habitat diversity decreased and riffles became filled with fine sediments (Jones, Helfman, Harper, & Bolstadt, 1999). As the length of the non-forested segments increased, overall fish abundance decreased, though the number of non-native species increased. Even in heavily forested areas, clearing a 1-3 km stretch of forested buffer was found to have substantial impacts on fish assemblages (Jones, Helfman, Harper, & Bolstadt, 1999).
A broad range of mammals, birds, reptiles and amphibians rely on riparian buffers for habitat. Riparian buffers are core habitat for many semi-aquatic and terrestrial ecotone species, such as salamanders, frogs, turtles, minks, beavers and otters, and these species require a buffer that is both long and wide. Long stretches of riparian buffer also serve as wildlife travel corridors. Many birds, such as herons, fishers, eagles, and ospreys, as well as some mammals, rely on forested buffers for both habitat and resting places. These birds hunt for fish in the water and nest in adjacent forests.
For buffers to provide adequate habitat for forest dependent songbirds, they must be wide. Several studies have shown that bird species richness increases in buffers that are at least 100 meters wide and that the presence of forest dependent songbirds decreases dramatically when buffers are less than 50 meters (Bongard, 2009). For more information on the importance of protecting species richness, see the guide Biodiversity.
Riparian buffers, especially forested buffers, absorb rainwater, which recharges ground water supplies and allows storm runoff to be released more slowly. This reduces the intensity and frequency of flooding as well as allows for more water flow in streams during dry periods.
The minimum width needed for an effective riparian buffer depends on the function you want the buffer to serve. For example, sediment can be physically filtered out of stormwater faster than dissolved nitrogen, which requires bacterial transformation to remove it. Thus, a narrower buffer would be needed to remove sediment than that needed to remove dissolved nitrogen. Scientific studies have shown that efficient buffer widths range from 10 feet for bank stabilization and stream shading to over 300 feet for wildlife habitat. (Hawes & Smith, 2005). Necessary widths will also vary depending on site conditions, such as soil type, slope and adjacent land use and other factors. (Hawes & Smith, 2005)
In Riparian Buffer Zones: Functions and Recommended Widths (Hawes and Smith, 2005), the authors summarize the results of scientific studies, identifying the buffer widths needed for a buffer to effectively serve particular functions; they report the following ranges:
Erosion/sediment control 30 feet to 98 feet
Nutrients 49 feet to 164 feet
Pesticides 49 feet to 328 feet
Biocontaminants 30 feet or more (e.g. fecal matter)
Wildlife 33 feet to 164 feet
Litter/debris 50 feet to 100 feet
Temperature 30 feet to 230 feet
Regarding terrestrial habitat, research suggests a range of 30 to 1,640 feet. However, because the habitat needs for terrestrial wildlife vary widely, the authors do not believe it is feasible to capture the needs of all species with a uniform buffer size. They recommend reviewing information about specific animals in the targeted area as well as land conservation work at adjacent and nearby lands.
Headwater streams are the smaller tributaries that carry water from the upper reaches of the watershed to the main channel of the river. They are rarely named and are often so small that it takes little effort to jump across them. While there is no universally accepted definition of headwaters, they are often defined as first and second order streams. A stream with no tributaries, recurring or perennial, is a first order stream. When two first-order streams come together, they form a second-order stream. The Stroud Research Center defines headwaters as “tributary streams, intermittent streams, and spring seeps” (Kaplan, Bott, Jackson, Newbold, & Sweeney, 2008).
Headwaters represent 50-70% of the total stream miles in the U.S. (Fritz, Johnson, & Walters, 2008). Nearly everyone in the United States has a headwater stream within a mile or two of their home, leaving headwaters close to human activities such as urbanization, dams and diversions, water withdrawals, point and non-point source pollution, deforestation, and agriculture (River Keeper, 2005). The small size of headwater streams, along with their integration into the landscape, makes them highly vulnerable to degradation (Kaplan et al., 2008).
Headwater streams are not as resilient as larger streams because they lack sufficient water flow to transport and dilute sediment and pollution (Kaplan et al., 2008). Forested buffers are needed to remove pollutants from stormwater before they reach the stream. The aquatic wildlife of headwaters are usually coldwater adapted (Kaplan et al., 2008), and therefore rely on the temperature moderation effects of riparian trees. Riparian buffers are essential to the provision of food for both the headwaters themselves, and the resulting downstream food web. Riparian vegetation provides up to 90% of the organic matter (food) necessary to support headwater stream communities (Cummins & Spengler, 1978).
Water quality, biodiversity, and ecological health of freshwater systems depend on the ecosystem services of healthy headwater streams (Kaplan et al., 2008). According to Lowe and Likens (2005),
There is no doubt that it is important to safeguard lowland sites, but it is difficult to see how any conservation action with a goal of protecting the long-term ecological integrity and ecosystem services of natural systems, whether aquatic or terrestrial, can succeed without a foundation of intact and functional headwaters.
Headwaters are the source of much of the water, gravel, wood, and nutrients that flow through the stream network and eventually to the ocean (USDA, 2008). Headwaters can help to keep sediment and pollutants out of the stream system’s lower reaches. (Kaplan et al., 2008).
Recycling organic carbon contained in the bodies of dead plants and animals is a crucial ecosystem service and is the basis for every food web on the planet (Meyer et al., 2003). In freshwater ecosystems, much of this recycling happens in small streams and wetlands (Meyer et al., 2003). This recycling process makes nutrients more biologically available to organisms downstream (Meyer et al., 2003). Headwater streams have been found to be significantly more efficient at breaking down the larger organic materials of dead plant and animals into nutrients usable to small animals, such as mayflies and caddis flies. The nutrients then work their way through the food web into larger animals downstream such as trout and birds. The processing of organic carbon in headwaters also prevents large amounts of organic material from being taken downstream, where the decomposition of large quantities could deplete dissolved oxygen levels and kill or harm aquatic life (Meyer et al., 2003).
Owing to favorable microclimate and availability of water, headwaters provide habitat for distinct assemblages of plants and animals (USDA, 2008). Hydrological conditions of many headwaters, which include running seasonally and drying out in the summer, periodically flowing underground, and frequent cascades and obstacles, lead to a lack of fish, which provides habitat that many amphibians can thrive in. Headwaters act as refugia for riverine species during specific life-history stages and critical periods of the year, such as warm summer months (Lowe & Likens, 2005).
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Castelle, A., and Johnson, A. (2000). Riparian Vegetation Effectiveness: Technical Bulletin No. 799. Research Triangle Park, NC: National Council for Air and Stream Improvement, Inc. 36pp.
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Dabney, S., Moore, M., and Locke, M. (2006). Integrated Management of In-Field, Edge-of-Field, and After-Field Buffers. Journal of the American Water Resources Association, 42(1),15-24. doi: 10.1111/j.1752-1688.2006.tb03819.x.
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Hawes, E. and Smith, M. (2005). Riparian Buffer Zones: Functions and Recommended Widths. Prepared for the Eightmile River Wild and Scenic Study Committee.
Jones, E., III, Helfman, G., Harper, J., and Bolstadt, P. (1999). Effects of Riparian Forest Removal on Fish Assemblages in Southern Appalachian Streams. Conservation Biology, 13(6). 1454-1465.
Kaplan, L., Bott, T., Jackson, J., Newbold, J.D., & Sweeney, B., 2008. Protecting Headwaters: The Scientific Basis for Safeguarding Stream and River Ecosystems. Stroud Water Research Center, Avondale, PA.
Klapproth, J. and Johnson, J. (2009). Understanding the Science Behind Riparian Forest Buffers: Effects on Plant and Animal Communities. Blacksburg, VA: Virginia Polytechnic Institute and State University. Publication 420-152
Lutz, L. (2006). Minebank Run Restoration Hits Pay Dirt in Reducing Nitrogen Loads. Chesapeake Bay Journal, 16(2). Retreived from http://www.bayjournal.com.
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Meyer, J.L., Kaplan, L.A., Newbold, D., Strayer, D., Woltemade, C.J., Zedler, J.B., Zedler, P.H. (2003). Where Rivers are Born: The Scientific Imperative for Defending Small Streams and Wetlands. Published by the Sierra Club and American Rivers.
Pitt, D. and Batzer, D. (2011). Woody Debris as a Resource for Aquatic Macroinvertebrates in Stream and River Habitats of the Southeastern United States: A Review. Proceedings of the 2011 Georgia Water Resources Conference, held April 11-14, 2011, at the University of Georgia.
Pollock, M., Beechie, T., Liermann, M., and Bigley, R. (2009). Stream Temperature Relationships to Forest Harvest in Western Washington. Journal of the American Water Resources Association, 45(1), 141-156. doi: 10.1111/j.1752-1688.2008.00266.x.
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Sweeney, B. (1992). Streamside Forests and the Physical, Chemical, and Trophic Characteristics of Piedmont Streams in Eastern North America. Water Science and Technology. 26 (12), 2653-2673.
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United States Environmental Protection Agency (EPA,1995). Linking Restoration Practices to Water Quality Parameters. Chapter 3 in Ecological Restoration: A Tool To Manage Stream Quality, Washington, D.C., U.S. EPA. EPA 841-F-95-007.
The Pennsylvania Land Trust Association published this guide with support from the William Penn Foundation, the Colcom Foundation and the Growing Greener Program of the Pennsylvania Department of Conservation and Natural Resources, Bureau of Recreation and Conservation.
© 2014 Pennsylvania Land Trust Association
Text may be excerpted and reproduced with acknowledgement of ConservationTools.org and the Pennsylvania Land Trust Association.